Aquatic
biomes occupy the largest part of the biosphere.
·
Ecologists
distinguish between freshwater and marine biomes on the basis of physical and
chemical differences.
°
Marine
biomes generally have salt concentrations that average 3%, while freshwater
biomes have salt concentrations of less than 1%.
·
Marine
biomes cover approximately 75% of the earth’s surface and have an enormous
effect on the biosphere.
°
The
evaporation of water from the oceans provides most of the planet’s rainfall.
°
Ocean
temperatures have a major effect on world climate and wind patterns.
°
Photosynthesis
by marine algae and photosynthetic bacteria produce a substantial proportion of
the world’s oxygen. Respiration by these organisms consumes huge amounts of
atmospheric carbon dioxide.
·
Freshwater
biomes are closely linked to the soils and biotic components of the terrestrial
biomes through which they pass.
°
The
pattern and speed of water flow and the surrounding climate are also important.
·
Most
aquatic biomes are physically and chemically stratified.
·
Light
is absorbed by the water and by photosynthetic organisms, so light intensity
decreases rapidly with depth.
°
There
is sufficient light for photosynthesis in the upper photic zone.
°
Very
little light penetrates to the lower aphotic
zone.
·
The
substrate at the bottom of an aquatic biome is the benthic zone.
°
This
zone is made up of sand and sediments and is occupied by communities of
organisms called benthos.
°
A
major food source for benthos is dead organic material or detritus, which rains down from the productive surface waters of
the photic zone.
·
Sunlight
warms surface waters, while deeper waters remain cold.
°
As
a result, water temperature in lakes is stratified, especially in summer and
winter.
°
In
the ocean and most lakes, a narrow stratum of rapid temperature change called a
thermocline separates the more
uniformly warm upper layer from more uniformly cold deeper waters.
·
In
aquatic biomes, community distribution is determined by depth of the water,
distance from shore, and open water versus bottom.
·
In
marine communities, phytoplankton, zooplankton, and many fish species live in
the relatively shallow photic zone.
·
The
aphotic zone contains little life, except for microorganisms and relatively
sparse populations of luminescent fishes and invertebrates.
·
The
major aquatic biomes include lakes, wetlands, streams, rivers, estuaries,
intertidal biomes, oceanic pelagic biomes, coral reefs, and marine benthic
biomes.
·
Freshwater
lakes vary greatly in oxygen and nutrient content.
°
Oligotrophic lakes are deep,
nutrient poor, oxygen rich, and contain little life.
°
Eutrophic lakes are shallow,
nutrient rich, and oxygen poor.
·
In
lakes, the littoral zone is the
shallow, well-lit water close to shore.
°
The
limnetic zone is the open surface
water.
·
Wetlands are areas covered
with sufficient water to support aquatic plants.
°
They
can be saturated or periodically flooded.
°
Wetlands
include marshes, bogs, and swamps.
°
They
are among the most productive biomes on Earth and are home to a diverse
community of invertebrates and birds.
°
Because
of the high organic production and decomposition in wetlands, their water and
soil are low in dissolved oxygen.
°
Wetlands
have a high capacity to filter dissolved nutrients and chemical pollutants.
°
Humans
have destroyed many wetlands, but some are now protected.
·
Streams and rivers are bodies of water moving
continuously in one direction.
°
Headwaters
are cold, clear, turbulent, and swift.
§ They carry little
sediment and relatively few mineral nutrients.
°
As
water travels downstream, it picks up O2 and nutrients on the way.
°
Nutrient
content is largely determined by the terrain and vegetation of the area.
§ Many streams and
rivers have been polluted by humans, degrading water quality and killing
aquatic organisms.
§ Damming and flood
control impairs the natural functioning of streams and rivers and threatens
migratory species such as salmon.
·
Estuaries are areas of
transition between river and sea.
°
The
salinity of these areas can vary greatly.
°
Estuaries
have complex flow patterns, with networks of tidal channels, islands, levees,
and mudflats.
°
They
support an abundance of fish and invertebrate species and are crucial feeding areas
for many species of waterfowl.
·
An
intertidal zone is a marine biome
that is periodically submerged and exposed by the tides.
°
The
upper intertidal zone experiences longer exposure to air and greater variation
in salinity and temperature than do the lower intertidal areas.
°
Many
organisms live only at a particular stratum in the intertidal.
·
The
oceanic pelagic biome is the open
blue water, mixed by wind-driven oceanic currents.
°
The
surface waters of temperate oceans turn over during fall through spring.
°
The
open ocean has high oxygen levels and low nutrient levels.
°
This
biome covers 70% of the Earth’s surface and has an average depth of 4,000
meters.
·
Coral reefs are limited to the
photic zone of stable tropic marine environments with high water clarity. They
are found at temperatures between 18°C and 30°C.
°
They
are formed by the calcium carbonate skeletons of coral animals.
°
Mutualistic
dinoflagellate algae live within the tissues of the corals.
°
Coral
reefs are home to a very diverse assortment of vertebrates and invertebrates.
°
Collecting
of coral skeletons and overfishing for food and the aquarium trade have reduced
populations of corals and reef fishes.
°
Global
warming and pollution contribute to large-scale coral mortality.
·
The
marine benthic zone consists of the
seafloor below the surface waters of the coastal or neritic zone and the offshore pelagic
zone.
°
Most
of the ocean’s benthic zone receives no sunlight.
°
Organisms
in the very deep abyssal zone are
adapted to continuous cold (about 3°C) and extremely high pressure.
°
Unique
assemblages of organisms are associated with deep-sea hydrothermal vents of volcanic origin on mid-ocean ridges.
§ The primary
producers in these communities are chemoautotrophic prokaryotes that obtain
energy by oxidizing H2S formed by a reaction of volcanically heated
water with dissolved sulfate (SO42−).
Climate largely determines the distribution and structure of
terrestrial biomes
·
Because
there are latitudinal patterns of climate over the Earth’s surface, there are
also latitudinal patterns of biome distribution.
·
A
climograph denotes the annual mean
temperature and precipitation of a region.
°
Temperature
and rainfall are well correlated with different terrestrial biomes, and each
biome has a characteristic climograph.
·
Most
terrestrial biomes are named for major physical or climatic features or for
their predominant vegetation.
·
Vertical
stratification is an important feature of terrestrial biomes.
°
The
canopy of the tropical rain forest
is the top layer, covering the low-tree stratum, shrub understory, ground
layer, litter layer, and root layer.
°
Grasslands
have a canopy formed by grass, a litter layer, and a root layer.
°
Stratification
of vegetation provides many different habitats for animals.
·
Terrestrial
biomes usually grade into each other without sharp boundaries. The area of
intergradation, called the ecotone,
may be narrow or wide.
·
The
species composition of any biome differs from location to location.
·
Biomes
are dynamic, and natural disturbance rather than stability tends to be the
rule.
°
Hurricanes
create openings for new species in tropical and temperate forests.
°
In
northern coniferous forests, snowfall may break branches and small trees,
producing gaps that allow deciduous species to grow.
°
As
a result, biomes exhibit patchiness, with several different communities
represented in any particular area.
·
In
many biomes, the dominant plants depend on periodic disturbance.
°
For
example, natural wildfires are an integral component of grasslands, savannas,
chaparral, and many coniferous forests.
·
Human
activity has radically altered the natural patterns of periodic physical
disturbance.
°
Fires
are now controlled for the sake of agricultural land use.
·
Humans
have altered much of the Earth’s surface, replacing original biomes with urban
or agricultural ones.
·
The
major terrestrial biomes include tropical forest, desert, savanna, chaparral,
temperate grassland, coniferous forest, temperate broadleaf forest, and tundra.
·
Tropical forests are found close to
the equator.
°
Tropical rain
forests
receive constant high amounts of rainfall (200 to 400 cm annually).
°
In
tropical dry forests, precipitation is highly seasonal.
°
In
both, air temperatures range between 25°C and 29°C year round.
°
Tropical
forests are stratified, and competition for light is intense.
°
Animal
diversity is higher in tropical forests than in any other terrestrial biome.
·
Deserts occur in a band
near 30° north and south latitudes and in the interior of continents.
°
Deserts
have low and highly variable rainfall, generally less than 30 cm per year.
°
Temperature
varies greatly seasonally and daily.
°
Desert
vegetation is usually sparse and includes succulents such as cacti and deeply
rooted shrubs.
°
Many
desert animals are nocturnal, so they can avoid the heat.
°
Desert
organisms display adaptations to allow them to resist or survive desiccation.
·
Savanna is found in
equatorial and subequatorial regions.
°
Rainfall
is seasonal, averaging 30–50 cm per year.
°
The
savanna is warm year-round, averaging 24–29°C with some seasonal variation.
°
Savanna
vegetation is grassland with scattered trees.
°
Large
herbivorous mammals are common inhabitants.
§ The dominant
herbivores are insects, especially termites.
°
Fire
is important in maintaining savanna biomes.
·
Chaparrals have highly
seasonal precipitation with mild, wet winters and dry, hot summers.
°
Annual
precipitation ranges from 30 to 50 cm.
°
Chaparral
is dominated by shrubs and small trees, with a high diversity of grasses and
herbs.
°
Plant
and animal diversity is high.
°
Adaptations
to fire and drought are common.
·
Temperate
grasslands
exhibit seasonal drought, occasional fires, and seasonal variation in
temperature.
°
Large
grazers and burrowing mammals are native to temperate grasslands.
°
Deep
fertile soils make temperate grasslands ideal for agriculture, especially for
growing grain.
°
Most
grassland in North America and Eurasia has been converted to farmland.
·
Coniferous forest, or taiga, is the largest terrestrial biome
on Earth.
°
Coniferous
forests have long, cold winters and short, wet summers.
°
The
conifers that inhabit these forests are adapted for snow and periodic drought.
°
Coniferous
forests are home to many birds and mammals.
°
These
forests are being logged at a very high rate and old-growth stands of conifers
may soon disappear.
·
Temperate broadleaf
forests
have very cold winters, hot summers, and considerable precipitation.
°
A
mature temperate broadleaf forest has distinct vertical layers, including a
closed canopy, one or two strata of understory trees, a shrub layer, and an
herbaceous layer.
°
The
dominant deciduous trees in Northern Hemisphere broadleaf forests drop their
leaves and become dormant in winter.
°
In
the Northern Hemisphere, many mammals in this biome hibernate in the winter,
while many bird species migrate to warmer climates.
°
Humans
have logged many temperate broadleaf forests around the world.
·
Tundra covers large areas
of the Arctic, up to 20% of the Earth’s land surface.
°
Alpine tundra is found on high
mountaintops at all latitudes, including the tropics.
§ The plant
communities in alpine and Arctic tundra are very similar.
°
The
Artic tundra winter is long and cold, while the summer is short and mild. The
growing season is very short.
°
Tundra
vegetation is mostly herbaceous, consisting of a mixture of lichens, mosses,
grasses, forbs, and dwarf shrubs and trees.
°
A
permanently frozen layer of permafrost
prevents water infiltration and restricts root growth.
°
Large
grazing musk oxen are resident in Arctic tundra, while caribou and reindeer are
migratory.
°
Migratory
birds use Arctic tundra extensively during the summer as nesting grounds.
Arctic tundra is sparsely settled by humans but has recently become the
focus of significant mineral and oil extraction
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